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The recovery of the airman follows separate search efforts by both the US and Iran.
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Posting to social media, US President Donald Trump said the rescued weapons systems officer had "sustained injuries" but would be "just fine", describing the rescue as "one of the most daring search and rescue operations in US history".
Following the downing of the aircraft on Friday, both the pilot and crew member ejected - with the pilot recovered shortly afterwards.
Following this, both the US and Iran scrambled to locate the missing crew member in a mountainous region of southwestern Iran.
The search presented a serious test for the US, with the possibility of a prisoner of war situation if Iran found the missing airman - something which would have likely involved him being used as part of propaganda material.
Iran's Islamic Revolutionary Guard Corps (IRGC) was said to have launched its own search for the missing American, reportedly using troops and locals, and offering them a reward of about $66,000 (£50,000) to capture him alive.
Videos on social media appeared to show hundreds of people heading to a mountainous area in south-western Iran to search for the missing US airman.
Details of the second airman being rescued emerged in US media late on Saturday night.
A short time later, Trump confirmed the successful mission on Truth Social, writing "WE GOT HIM!", adding the crew member was a "respected colonel".
The US rescue operation was described to the BBC as "huge" by one person familiar with the details.
The BBC understands that there was an engagement between the US and Iranian forces during the rescue, and that the pilot may have been injured during his initial ejection from the aircraft.
Tasnim, the semi-official news agency associated with the IRGC, said five Iranians were killed during the operation.
Trump said dozens of aircraft had been involved in the operation, with the White House not giving an update after the pilot was recovered on Friday to protect the ongoing rescue.
Earlier reports suggested the missing crew member might be in
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